| bole | habit | large trees 30+ m or small trees or shrubs |
| bole | buttresses | absent |
| bole | spines | absent |
| bole | bark_outer_color | yellow or gray or brown |
| bole | bark_peeling | not peeling |
| bole | slash_lamination | absent |
| bole | slash_odor | absent |
| bole | exudate | absent |
| twig | terminal_buds_enclosed_by_leaves | no |
| twig | clumped_hairs | absent |
| twig | complex_hairs | absent |
| twig | stinging_hairs | absent |
| twig | mature_twig_indumentum | absent |
| twig | twig_cross_section | terete |
| twig | leaf_location | clustered at twig end |
| twig | leaf_insertion | opposite |
| twig | leaf_plane | two planes |
| stipule | stipules | present |
| stipule | stipule_freedom | free |
| stipule | stipule_type | lateral |
| stipule | stipule_on_twig | not encirlcing twig |
| stipule | stipule_form | deltoid |
| stipule | stipule_fringe | absent |
| stipule | stipule_size | minute |
| stipule | stipule_persistence | deciduous |
| leaf | leaf_scalelike_or_needlelike | none |
| leaf | leaf_complexity | simple |
| leaf | petiole_length | less than a quarter or quarter to half |
| leaf | petiole_winged | not winged |
| leaf | petiole_attachment | base of leaf |
| leaf | petiole_base_swelling | not swollen |
| leaf | petiole_apex_swelling | not swollen |
| leaf | petiole_cross_section | flattened or grooved |
| leaf | petiole_wrinkling | longitundinal |
| leaf | leaf_similarity | homophyllus |
| leaf | leaf_texture | coriaceous |
| leaf | leaf_base | symmetrical |
| leaf | leaf_dissection | not dissected |
| leaf | leaf_margins | toothed |
| leaf | margin_plane | flat |
| leaf | leaf_base_shape | tapered |
| leaf | leaf_apex_shape | retuse or rounded or acute |
| leaf | upper_leaf_surface | matte |
| leaf | lower_leaf_surface | matte |
| leaf | primary_venation | pinnate |
| leaf | primary_vein_above | flat or sunken |
| leaf | secondary_vein_spacing | open |
| leaf | secondaries_above | flat |
| leaf | secondary_vein_course | mostly parallel |
| leaf | secondary_vein_angle | medium |
| leaf | secondary_vein_termination | intramarginal |
| leaf | secondary_vein_branching | not branched or branched |
| leaf | intersecondary_veins | absent |
| leaf | tertiary_veins_above | not visible |
| leaf | tertiary_vein_form | weakly percurrent or reticulate |
| leaf | direction_of_percurrence | perpendicular to midrib or parallel to midrib |
| leaf | ramified_branching | absent |
| leaf | domatia | absent |
| leaf | leaf_glands | absent |
| leaf | marginal_and_petiolar_glands | absent or laminar margin |
| leaf | hairs_on_mature_leaf | absent |
| inflor | inflorescence | multiple |
| inflor | inflorescence_position | axillary |
| inflor | inflorescence_axis | branched |
| inflor | specialized_inflorescence_units | absent |
| flower | numerous_persistant_floral_bracts | absent |
| flower | floral_sexuality | bisexual |
| flower | peduncle | present |
| flower | flower_size | to about 0.5 cm |
| flower | floral_symmetry | actinomorphic |
| flower | floral_column | absent |
| flower | hypanthium | absent |
| flower | disk | present |
| flower | disk_position | stamens outside (intrastaminal) or stamens on disk |
| flower | perianth | present |
| flower | perianth_type | K+C |
| flower | sepal_number | 4-5 |
| flower | sepal_freedom | free or partially joined |
| flower | sepal_persistence | caducous |
| flower | sepal_accrescence | not accrescent |
| flower | epicalyx | absent |
| flower | petal_presence | present |
| flower | petal_number | 4-5 |
| flower | petal_fusion | free at base |
| flower | petal_texture | not fleshy |
| flower | petal_persistence | deciduous |
| flower | petal_stalks | sessile |
| flower | petal_margin | entire |
| male | stamen_number | 4-5 |
| male | stamen_filaments | present |
| male | stamen_fascicling | all free |
| male | stamen_adnation | free |
| male | stamen_size | equal sizes |
| male | stamen_mery | isomerous |
| male | isomerous_stamen_position | oppositisepalous |
| male | anther_locule_number | 2 |
| male | anther_fixation | dorsifixed |
| male | anther_adornment | not awned |
| male | anther_dehiscence | long slits |
| female | ovary_position | superior |
| female | carpel_arrangement | syncarpous |
| female | carpel_number | 2 |
| female | locule_number | 2 |
| female | style_presence | absent or solitary including joined |
| female | stigma_lobes | 2 |
| female | stigma_shape | not distinctive |
| fruit | common_fruit_type | drupe |
| fruit | fruit_fleshiness | not fleshy or fleshy |
| fruit | fruit_aggregation | not multiple or aggregate |
| fruit | fruit_dehiscence | indehiscent |
| fruit | fruit_accessory_tissue | no fleshy perianth parts |
| fruit | fruit_wings | absent |
| seed | seed_number_and_multiple | 1-2 |
| seed | seed_size | less than 5 mm |
| seed | seed_wings | absent |
| seed | seed_ruminations | seed uniform |
| seed | seed_aril | absent |
| seed | seed_endosperm | present |
| info | field_characters | leaves decussate; leaves sometimes slightly toothed apically; ovary 2-celled; style very short or apparently absent; fruit a drupe |
| info | scoring_decisions | leaf margin changed to from all types to toothed. All herbarium specimens (not just Borneo) had teeth; usually along the whole margin length; not just near the apex. |
